The X-ray and infrared pictures reveal that at some point, three of the figures were clothed. Bellini lowered their necklines, presumably at the request of Duke Alfonso, to make the scene more "classical." Please touch numbers to clothe figures.
Who unclothed the nymphs? Was it Bellini? Probably. Bellini used his fingertips to meld the wet paint colors together in a characteristic manneras on the edge of the eroticized nymph's blue dress. And the other changes? X-ray and infrared do reveal other changes to the figures, such as moved hands or added attributes like Neptune's trident. If Bellini eroticized the figures, it is likely he (and not Titian) also made these other changes.